pcray1231 wrote:
I'd guess heat. At least for evening hatches.
Heat, i.e. water temperature, is the biggest thing regarding the timing of the hatch. First you need mature nymphs, so that's the total "temperature days", meaning over the entire year, not just during the season in which they hatch. Then, having that, water temperature offers a "trigger", so when it reaches X degrees the bugs that are mature start to hatch en masse.
That's why you typically have larger individuals within a species hatching first. Those nymphs were mature long before the trigger came. Later in the hatch, the limiting factor is the maturity of the nymphs, so they hatch as soon as they are able.
Diurnal cycles, and perhaps moon phase, weather (cloud cover), pressure, air temperature, are potentially important as well, most of which are harder to prove.
But heat apparently isn't how they navigate AFTER hatching. While mating, it's an attraction to light (at dusk, they come out of trees in the dark woods and hover over streams with an open canopy, where it's brighter). After mating, and it's time to locate the stream for egg laying, it's horizontally polarized light that attracts them. Starlight or moonlight off of a dark water surface will do it. But off of a dark smooth road surface will as well. And a streetlight over top a dark smooth road surface offers attraction at both stages.